Successfully ruled out A, B, and C. I was stuck between D and E and honestly, I had no idea at the time so I guessed incorrectly.
Looking back on it now, perhaps D is incorrect because fossils in the upland found together with products of an arid landscape would go against his hypothesis because it would give some credibility to the idea that the uplands also turned desert-like?
As for E, i have no idea what they are talking about really...is E correct because it gives proof that the lowlands were desert-like during the ice age, yet not as desert-like as they are now?